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Peer-reviewed publications

Twomey, E, and J.L. Brown. 2008.  A partial revision of the Ameerega hahneli complex and a new cryptic species from the East- Andean versant of Central Peru. Zootaxa 1757: 49-65.

Twomey, E, and J.L. Brown. 2008. Spotted poison frogs: Rediscovery of a lost species and new genus (Anura: Dendrobatiade) from Northwestern Peru. Herpetologica 64: 121-137. PDF

Brown, J.L., E. Twomey, V. Morales, and K. Summers.  Phytotelm size in relation to parental care and mating strategies in two species of Peruvian poison frogs. Behaviour (in press).

Twomey, E., V. Morales, and K. Summers. Evaluating condition-specific and asymmetric competition in a species-distribution context. Oikos (in press).


Roberts, J.L., J.L. Brown, R. Von May, R. Schulte, and K. Summers. 2006. Phylogenetic relationships among  members of the genus Dendrobates: A molecular perspective.  Herpetological Journal 16: 377-385. PDF

Roberts, J.L., J.L. Brown, R. Schulte, W. Arizabal, and K. Summers. 2006. Genetic divergence and speciation in lowland and montane Peruvian poison frogs.  Journal of Molecular Phylogenetics and Evolution. 41:149-164. PDF

Roberts, J.L., J.L. Brown, R. Schulte, W. Arizabal, and K. Summers. 2006. Rapid diversification of colouration among populations of a poison frog isolated on sky peninsulas in the central cordilleras of Peru. Journal of Biogeography 34: 417-426. PDF

Brown, J.L., R. Schulte, and K. Summers. 2006. A new species of Dendrobates (Anura: Dendrobatidae) from the Amazonian Lowlands in Peru.  Zootaxa 1152: 45-58  PDF

Twomey, E, and J.L. Brown. 2005. Range extension: Cochranella respendens Herpetological Review, 1 page.

Brown, J.L., and A.L. Vaughan. 2003) Species note: Arboreal Celestus. A field observation. Herpetological Review, 1 page. PDF

Wisenden, B.D., K.A. Vollbrecht, and J.L. Brown. 2002. Is there a fish alarm cue? Affirming evidence from the wild study.  Animal Behavior 67: 59-67 PDF

Other Publications

Brown JL, and E. Twomey (in press).   Ranitomeya imitator, R. fantastica, Ameerega bassleri.  Species accounts in the book: Andean Poison Frogs. Conservation International.  

Brown, J.L., and K. Siu Ting (in press).   Ranitomeya uakarii and Ameerega cainrachi. Species accounts in the book: Andean Poison Frogs. Conservation International.  

Twomey, E., J.L Brown, and J. Yeager (in press).  Excidobates captivus. Species account in the book: Andean Poison Frogs. Conservation International.  

Pepper, M., E. Twomey, and J.L. Brown. 2007. The smuggling crisis. Leaf Litter Magazine.  Tree Walkers International. Spring 2007 Vol. 1.1 pp 5-7. PDF

Brown, J.L. 2007.  Peru’s Tiny Gems. Photograph.  National Geographic Magazine. April 2007. PDF

Copyright (c) Evan Twomey and Jason Brown

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