Publications

2016
Twomey, E., Vestergaard, J. S., Venegas, P. J., & Summers, K. (2016). Mimetic Divergence and the Speciation Continuum in the Mimic Poison Frog Ranitomeya imitator. The American Naturalist, 187(2), 205–224.

2015
Twomey, E., Mayer, M., & Summers, K. (2015). Intraspecific Call Variation in the Mimic Poison Frog Ranitomeya imitator. Herpetologica, 71(4), 252–259.

Vestergaard, J. S., Twomey, E., Larsen, R., Summers, K., & Nielsen, R. (2015). Number of genes controlling a quantitative trait in a hybrid zone of the aposematic frog Ranitomeya imitator. Proceedings of the Royal Society B: Biological Sciences, 282, 1–10.

2014
Mayer, M., Schulte, L. M., Twomey, E., & Lötters, S. (2014). Do male poison frogs respond to modified calls of a Müllerian mimic? Animal Behaviour, 89, 45–51.

Twomey, E., Vestergaard, J. S., & Summers, K. (2014). Reproductive isolation related to mimetic divergence in the poison frog Ranitomeya imitator. Nature Communications, 5(4749), 1–8.

2013

Brown, J. L. (2013a). Neotropical poison frogs: evolution’s guide to parenting, fashion and communication in a dynamic world. Evolutionary Ecology, 27(4), 655–659.

Brown, J. L. (2013b). The evolution of parental care, aposematism and color diversity in Neotropical poison frogs. Evolutionary Ecology, 27(4), 825–829.

Gehara, M., Summers, K., & Brown, J. L. (2013). Population expansion, isolation and selection: novel insights on the evolution of color diversity in the strawberry poison frog. Evolutionary Ecology, 27(4), 797–824.

Twomey, E., Yeager, J., Brown, J. L., Morales, V., Cummings, M., & Summers, K. (2013). Phenotypic and genetic divergence among poison frog populations in a mimetic radiation. PloS One, 8(2), e55443.

2012

Yeager, J., Brown, J. L., Morales, V., Cummings, M., & Summers, K. (2012). Testing for selection on color and pattern in a mimetic radiation. Current Zoology, 58(4), 667–675.

2011

Brown, J.L., E. Twomey, A. Amezquita, M. Barbosa de Souza, J.P. Caldwell, S. Lötters, R. von May, P.R. Melo Sampaio, D. Mejia-Vargas, P. Perez-Peña, M. Pepper, E.H. Poelman, M. Sanchez-Rodriguez, & K. Summers. 2011. A taxonomic revision of the Neotropical poison frog genus Ranitomeya (Amphibia: Dendrobatidae). Zootaxa, 3083: 1-120.

2010

Brown, J.L., V. Morales, and K. Summers. 2010. A key ecological trait drove the evolution of biparental care and monogamy in an amphibian. The American Naturalist 175: 436-446.

Brown, J.L., M.E. Maan, M.E. Cummings, and K. Summers. 2010. Evidence for selection on coloration in a Panamanian poison frog: a coalescent-based approach. Journal of Biogeography 37: 891-901.

Perez-Peña, P.E., G. Chavez, E. Twomey, and J.L. Brown. 2010. Two new species of Ranitomeya (Anura: Dendrobatidae) from eastern Amazonian Peru. Zootaxa 2439: 1-23. | PDF

2009

Twomey, E., and J.L. Brown. 2009. Another new species of Ranitomeya (Anura: Dendrobatidae) from Amazonian Colombia. Zootaxa 2302: 48-60. | PDF

Brown, J.L., and E. Twomey. 2009. Complicated histories: three new species of poison frogs of the genus Ameerega (Anura: Dendrobatidae) from north-central Peru. Zootaxa 2049: 1-38. |PDF

Brown, J.L., V. Morales, and K. Summers. 2009. An experimental demonstration of tactical parasitism of conspecific reproduction via larval cannibalism in Peruvian poison frogs. Biology Letters 5: 148-151. | PDF

Brown, J.L., V. Morales, and K. Summers. 2009. Home range size and location in relation to reproductive resources in poison frogs: a Monte Carlo approach using GIS data. Animal Behaviour77: 547-554. | PDF

2008

Brown, J.L., V. Morales, and K. Summers. 2008.  The evolution of parental care, habitat selection and larval life history in Peruvian poison frogs. Journal of Evolutionary Biology 21: 1534-1543. | PDF

Brown, J.L., E. Twomey, M. Pepper, and M. Sanchez Rodriguez. 2008. A revision of the Ranitomeya fantastica species complex and two new species of poison frogs (Anura: Dendrobatidae) from the Rio Huallaga drainage in central Peru. Zootaxa 1832: 1-24. | PDF

Brown, J.L., E. Twomey, V. Morales, and K. Summers. 2008. Phytotelm size in relation to parental care and mating strategies in two species of Peruvian poison frogs. Behaviour 145: 1139-1165. | PDF

Twomey, E., V. Morales, and K. Summers. 2008. Evaluating condition-specific and asymmetric competition in a species distribution context. Oikos 117: 1175-1184. | PDF

Twomey, E., and J.L. Brown. 2008. Spotted poison frogs: rediscovery of a lost species and a new genus. Herpetologica 64: 121-137. | PDF

Twomey, E., and J.L. Brown. 2008. A partial revision of the Ameerega hahneli complex (Anura: Dendrobatidae) and a new cryptic species from the east-Andean versant of central Peru.Zootaxa 1757: 49-65. | PDF

Twomey, E., V. Morales, and K. Summers. 2008. Effects of kinship on intraspecific competition in larvae of the poison frog Ameerega bassleri (Anura: Dendrobatidae). Phyllomedusa 7: 121-126. | PDF

von May, R., Catenazzi, A., Angulo, A., Brown, J.L., Carrillo, J., Chávez, G., Córdova, J.H., Curo, A., Delgado, A., Enciso, M.A., Gutiérrez, R., Lehr, E., Martínez, J.L., Medina-Müller, M., Miranda, A., Neira, D.R., Ochoa, J.A., Quiroz, A.J., Rodríguez, D.A., Rodríguez, L.O., Salas, A.W., Seimon, T., Seimon, A., Siu-Ting, K., Suárez, J., Torres, J., and Twomey, E. 2008. Current state of conservation knowledge of threatened amphibian species in Peru. Tropical Conservation Science 1: 376-396. | PDF

2007

Roberts, J.L., Brown, J.L., Schulte, R.,  Arizabal, W, and Summers, K. 2007. Rapid diversification of coloration among populations of a poison frog isolated on sky peninsulas in the central cordilleras of Peru. Journal of Biogeography 34: 417-426. | PDF

2006

Roberts, J.L., J. L. Brown, R. von May, W. Arizabal, A. Presar, R. Symula, R. Schulte, and K. Summers. 2006. Phylogenetic relationships among poison frogs of the genus Dendrobates(Dendrobatidae): A molecular perspective and increased taxon sampling. The Herpetological Journal 16: 377-385. | PDF

Roberts, J.L., Brown, J.L., Arizabal, W., von May, R.,, Schulte, R., & Summers, K.  2006. Genetic divergence and speciation in lowland and montane Peruvian poison frogs.  Molecular Phylogenetics and Evolution 41: 149-164. | PDF

Brown, J.L., Schulte, R. & Summers, K. 2006. A new species of Dendrobates (Anura: Dendrobatidae) from the Amazonian lowlands in Peru.  Zootaxa 1152: 45-58. | PDF

Other Publications

Brown JL, Twomey E (in press). Ranitomeya imitator, R. fantastica, Ameerega bassleri.  Species accounts in the book: Andean Poison Frogs. Conservation International.

Brown JL, Siu Ting K (in press). Ranitomeya uakarii and Ameerega cainrachi. Species accounts in the book: Andean Poison Frogs. Conservation International.

Twomey E, Brown JL, Yeager J (in press). Excidobates captivus. Species account in the book: Andean Poison Frogs. Conservation International.